That is great that she has found comfort in making friends with the boys instead. She sounds like she has a lot of confidence! I think it is important to be aware of what is going on for your child although it sounds like you have this covered. You mentioned that your daughter is quite honest and open with you which is amazing!
What activities do you do when you spend time with your daughter? This can still provide a sense of friendship and help foster a healthy and beautiful relationship between you which will act as a model for other relationships.